Whilst up high a model may not shoot weapons at troops on the table, cast spells on to the table, or do anything to troops on the table. Similarly, a model that is up high cannot be shot at by troops on the table or have spells cast upon it by wizards on the table. The process of flying high puts you above and beyond the battlefield.
A model that is up high may attack an enemy model that is also up high. These attacks represent glancing blows inflicted while the combatants swoop and climb, circling round each other and 'dog fighting' in the sky. Glancing blows are worked out differently from ordinary combat. All combat fought whilst flying up high is struck in this way; ordinary combat is never fought between models that are up high.